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124 CHAPTER 6 Input 1 Input 2 X?(t), X?(t) 0.5 0.5 Input X? Input X? w? w? (X? OR X?) AND Y 0.5 0 0.5 3 Output (a) 0.5 0.5 Input X? Input X? 4 6 Y(t) X?(t), X?(t) 0.5 0 0 2 0.5 0 Time (c) Time (b) 5 FIGURE 6.22 Dynamics in a model of a C. elegans neuronal multi-input FFL following pulses of input stimuli. (a) A two-input FFL. The input functions and activation thresholds are shown. (b) Dynamics of the two-input FFL in response to well-separated input pulses that stimulate X? and X?, followed by a persistent X? stimulus. The separated pulses do not activate Z. (c) Dynamics with a brief X? stimulus followed closely by a short X? stimulus. The dashed horizontal line corresponds to the activation threshold for Y. The brief pulses are able to activate Z. Relaxation rate is ? = 1, and w? = w? = w'? = w'? = 0.5, w'? = 0.4. (From Kashtan et al., 2004.)

          124 CHAPTER 6
Input 1
Input 2
X?(t), X?(t)
0.5
0.5
Input X?
Input X?
w? w?
(X? OR X?)
AND
Y
0.5
0
0.5
3
Output
(a)
0.5
0.5
Input X?
Input X?
4
6
Y(t)
X?(t), X?(t)
0.5
0
0
2
0.5
0
Time
(c)
Time
(b)
5
FIGURE 6.22 Dynamics in a model of a C. elegans neuronal multi-input FFL following pulses of input
stimuli. (a) A two-input FFL. The input functions and activation thresholds are shown. (b) Dynamics of the
two-input FFL in response to well-separated input pulses that stimulate X? and X?, followed by a persistent
X? stimulus. The separated pulses do not activate Z. (c) Dynamics with a brief X? stimulus followed closely
by a short X? stimulus. The dashed horizontal line corresponds to the activation threshold for Y. The brief
pulses are able to activate Z. Relaxation rate is ? = 1, and w? = w? = w'? = w'? = 0.5, w'? = 0.4. (From Kashtan
et al., 2004.)

124 CHAPTER 6
Input 1
Input 2
X?(t), X?(t)
0.5
0.5
Input X?
Input X?
w? w?
(X? OR X?)
AND
Y
0.5
0
0.5
3
Output
(a)
0.5
0.5
Input X?
Input X?
4
6
Y(t)
X?(t), X?(t)
0.5
0
0
2
0.5
0
Time
(c)
Time
(b)
5
FIGURE 6.22 Dynamics in a model of a C. elegans neuronal multi-input FFL following pulses of input
stimuli. (a) A two-input FFL. The input functions and activation thresholds are shown. (b) Dynamics of the
two-input FFL in response to well-separated input pulses that stimulate X? and X?, followed by a persistent
X? stimulus. The separated pulses do not activate Z. (c) Dynamics with a brief X? stimulus followed closely
by a short X? stimulus. The dashed horizontal line corresponds to the activation threshold for Y. The brief
pulses are able to activate Z. Relaxation rate is ? = 1, and w? = w? = w'? = w'? = 0.5, w'? = 0.4. (From Kashtan
et al., 2004.)

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