Which statement about the gas exchange system of birds is not true?
a. Respiratory gases are not exchanged in the air sacs.
b. A bird can achieve more complete exchange of $\mathrm{O}_{2}$ from air to blood than humans can.
c. Air passes through birds' lungs in only one direction.
d. The gas exchange surfaces in bird lungs are the alveoli.
e. A breath of air remains in the system for two breathing cycles.